Cannabis Sativa L.

A Critical Review on Cannabinoids, Legislation, and Analytical Methods

Cannabinoids Legislation Analytical Methods

Introduction: The Versatile Plant

Cannabis sativa L., one of the most fascinating and controversial plants in human history, is gaining a prominent place in the global scientific and regulatory landscape. With a history dating back more than 4000 years of medicinal use in China, where the legendary Emperor Shen Nung documented its therapeutic applications around 2700 BC 1 , cannabis is now emerging from the shadows of prohibition to become the focus of innovative scientific research and a complex international regulatory puzzle.

Historical Significance

Cannabis has been used for medicinal purposes for over 4000 years, with documented use in ancient China 1 .

Modern Relevance

Today, cannabis is at the center of scientific research with over 500 identified compounds, about 100 of which are unique cannabinoids 1 .

In recent years, we have witnessed a quiet revolution in how cannabis is perceived, studied, and regulated. While laboratories worldwide examine the more than 500 compounds already identified in this unique plant, governments face the challenge of creating legal frameworks that balance the plant's therapeutic potential with the risks associated with its recreational use.

The Cannabis Plant and Its Key Compounds

Cannabinoids: The Main Actives

At the heart of scientific interest in cannabis are cannabinoids, unique terpenophenolic compounds of this plant . These molecules, which act primarily through our body's endocannabinoid system - a complex signaling network discovered only in the late 20th century - are responsible for the plant's therapeutic and psychoactive effects 1 .

Δ9-tetrahydrocannabinol (THC)

The main psychoactive component of cannabis, responsible for the characteristic "high" sensation. THC acts as a partial agonist of CB1 receptors in the brain 5 , triggering a cascade of effects that include not only psychoactivity but also pain relief, nausea reduction, and appetite stimulation 1 5 .

Cannabidiol (CBD)

A non-psychoactive cannabinoid with a remarkably diverse pharmacological profile. CBD has demonstrated anticonvulsant, anxiolytic, anti-inflammatory, and neuroprotective properties 5 . Unlike THC, CBD has little affinity for classical cannabinoid receptors, acting through other mechanisms including modulation of serotonin receptors and inhibition of endocannabinoid reuptake 1 5 .

Terpenes: More Than Just Aroma

In addition to cannabinoids, cannabis produces an impressive variety of terpenes - aromatic compounds that give the plant its distinctive odoriferous character 2 . These compounds, also found in other aromatic plants such as pines and citrus, are not merely ornamental; they actively contribute to the therapeutic effects of cannabis through what has been called the "entourage effect" 2 .

This phenomenon describes the synergistic action between cannabinoids and terpenes, where the plant's non-psychoactive compounds, including terpenes and flavonoids, modulate and complement the effects of the main cannabinoids.

  • Myrcene: A terpene also found in hops that appears to enhance the sedative effects of THC
  • Pinene: May counteract some of the memory deficits associated with THC 2
  • Limonene: Common in citrus varieties, contributes to mood elevation
Key Insight

It's crucial to understand that these cannabinoids exist in the plant mainly in their acidic forms (THCA and CBDA), which convert to neutral forms (THC and CBD) through a decarboxylation process, typically induced by heat . This transformation is essential for activating THC's psychoactive properties, explaining why raw cannabis does not produce the same effects as when heated.

Cannabis on the World Stage

The global legal landscape of cannabis is undergoing a profound and accelerated transformation. What was once a marginal issue has become a complex regulatory puzzle with significant economic and public health implications.

Country Medical Use Status Adult Use Status Relevant Notes
Germany Legalized & Expanding Legalized (since 2024) Adults can cultivate up to 3 plants; imports jumped from 32 tons (2023) to >70 tons (2024) 4
Portugal Legalized (since 2018) Decriminalized Largest medical exporter in Europe (32 tons in 2024), but local access limited by high prices 4
Spain Under Regulation Social Clubs Tolerated Royal Decree project submitted to EC in Jan 2025; >600 social clubs operating in de facto tolerance 4
Netherlands Legalized Experimentation Ongoing "Closed Coffee Shop Chain Experiment" in 75 municipalities; 7 licensed growers 4
United Kingdom Medical Prescription (NHS & private) Illegal Rapidly growing medical market; annual sales >200 million GBP 4
Poland Legal, but Restricted Illegal Medical consumption increased 224% in 2023; private telemedicine banned in Nov 2024 4
USA Varies by State Varies by State Federal rescheduling process stopped in April 2025 4
Economic Impact

The European medicinal cannabis market is estimated to grow from USD 2.59 billion in 2024 to USD 12.65 billion by 2033, representing a compound annual growth rate (CAGR) of 18.33% 4 .

18.33%

Projected CAGR for European Medicinal Cannabis Market (2024-2033)

How Scientists Analyze Cannabis

Gas Chromatography (GC)

Gas chromatography is one of the most established methods for the analysis of cannabinoids and terpenes . This technique involves vaporizing the sample and separating its components in a capillary column under high temperatures.

Limitation: The high temperatures required for analysis cause decarboxylation of acidic cannabinoids (THCA, CBDA), converting them to their neutral forms (THC, CBD) 2 .

GC is typically coupled with flame ionization detectors (FID) for routine quantification, or mass spectrometers (MS) for unambiguous compound identification .

High Performance Liquid Chromatography (HPLC)

Liquid chromatography has emerged as a powerful alternative to GC, particularly for cannabinoid analysis. The main advantage of HPLC lies in its ability to quantify both acidic and neutral forms of cannabinoids without the need for derivation, since separation occurs at lower temperatures that do not cause decarboxylation .

Modern HPLC systems generally use C18 stationary phase columns and mobile phases that combine methanol or acetonitrile with water acidified with formic acid .

When coupled with tandem mass spectrometry (MS/MS), HPLC becomes the gold standard for cannabinoid quantification in complex matrices, offering exceptional sensitivity and selectivity .
Parameter Gas Chromatography (GC) Liquid Chromatography (HPLC)
Sample Preparation May require derivation for acids Generally straightforward
Analysis Temperature High (causes decarboxylation) Ambient or moderate
Acidic Cannabinoid Forms Not directly quantified Directly quantified
Neutral Cannabinoid Forms Directly quantified Directly quantified
Terpene Analysis Excellent Limited
Sensitivity High Very high (with MS/MS)
Instrumental Cost Moderate High (for LC-MS/MS)

A Closer Look at a Key Experiment

Development of a GC Method for Simultaneous Analysis

One of the most significant analytical challenges in the field of cannabis science is the simultaneous analysis of terpenes and cannabinoids, compounds with drastically different physicochemical properties. An innovative study published in 2020 sought to overcome this challenge through the development of a unified gas chromatography method for the quantification of both groups of compounds in plant samples and their extracts 2 .

The research team faced two main obstacles: the large differences in polarity and volatility between terpenes and cannabinoids, and the need for exhaustive decarboxylation of the acidic forms of cannabinoids without degradation of volatile terpenes 2 .

Methodology: Step by Step
Optimized Extraction

Researchers tested several solvents before selecting acetone as the ideal extraction solvent, offering an effective compromise between the recovery of polar terpenes and lipophilic cannabinoids. Any solvent evaporation step after extraction was intentionally avoided to prevent significant losses of volatile terpenes 2 .

Sample-Solvent Ratio

A ratio of 1:17 (300 mg sample to 5 mL solvent) was established as the ideal balance between extraction efficiency and adequate terpene concentrations for quantification without the need for concentration steps 2 .

Chromatographic Separation

To solve the critical problem of overlap between cannabichromene (CBC) and cannabidiol (CBD), researchers selected a column with a 50% phenyl 50% dimethylpolysiloxane stationary phase, significantly more polar than the conventional 5% phenyl columns typically used in cannabinoid analysis 2 .

Injection and Temperature Conditions

The temperature program was carefully optimized to accommodate the wide range of volatilities, from highly volatile monoterpenes to heavy, low-volatility cannabinoids.

Results and Scientific Impact

The developed method demonstrated robust performance for both analyte groups. The detection limits ranged between 120-260 ng/mL for terpenes and 660-860 ng/mL for cannabinoids 2 . Parallel validation with established HPLC methods confirmed that the results for cannabinoids were comparable, validating the method's accuracy for routine applications.

Compound Detection Limit (ng/mL) Linear Range (μg/mL) Notes
β-myrcene 120 1-100 Predominant terpene in indica varieties
α-pinene 150 1-100 Contributes to "spicy" aroma
limonene 180 1-100 Common in citrus varieties
THC 750 10-1500 Main psychoactive
CBD 660 10-1500 Non-psychoactive, therapeutic
CBC 820 10-1500 Minor cannabinoid
This approach represents a significant methodological advance by allowing complete characterization of the chemical profile of cannabis in a single analysis, with minimal sample preparation and without resorting to expensive multidimensional chromatography equipment.

The Scientist's Toolkit

Modern cannabis analysis requires a variety of specialized reagents, solvents and reference materials to ensure accurate and reproducible results.

Item Function Specific Application Considerations
Certified Cannabinoid Standards Calibration and quantification GC, HPLC, LC-MS/MS Must include acidic and neutral forms; critical for accuracy
HPLC Grade Solvents Extraction and mobile phase Sample preparation, chromatography Low UV-abs for UV detection; free of interferences
Acetonitrile and Methanol Mobile phases HPLC, LC-MS High purity grade to avoid contamination
Acetone Extraction solvent Simultaneous extraction of terpenes and cannabinoids Selected as ideal compromise 2
Derivatizing Agents (e.g., BSTFA) Protection of functional groups GC for acidic forms Stabilizes acidic cannabinoids against decarboxylation
Chromatographic Columns Compound separation GC, HPLC 50% phenyl phase for improved separation 2
Acidified Mobile Phases Improve separation and peak shape HPLC 0.1% formic acid common
Quality Control Materials Method validation Quality assurance Matrix reference materials
2,4-dimethyl-9H-pyrido[2,3-b]indoleBench ChemicalsBench Chemicals
4,5,5-trifluoropent-4-enoic AcidBench ChemicalsBench Chemicals
2,4-bis(2-phenylpropan-2-yl)phenolBench ChemicalsBench Chemicals
2-Hexynyl-NECABench ChemicalsBench Chemicals
4-hydroxy-N-methylprolineBench ChemicalsBench Chemicals

Conclusion and Future Perspectives

Cannabis sativa L. has traveled a remarkable path from its ancient roots to becoming a focus of modern scientific and regulatory innovation. As research continues to unravel the mysteries of this complex plant, we are presented with new therapeutic applications and analytical challenges.

Analytical Techniques

Will continue to evolve, with multidimensional chromatography and high-resolution mass spectrometry offering increasingly detailed views of the plant's chemical complexity.

Clinical Research

Will deepen into the synergy between cannabinoids and terpenes, potentially leading to specific formulations for particular medical conditions.

Regulatory Frameworks

Will face the challenge of harmonizing rigorous quality control with equitable access for patients.

The Path Forward

As cannabis continues to transition from controlled substance to agricultural commodity and pharmaceutical, collaboration between scientists, regulators, healthcare professionals, and patients will be crucial to realizing the full potential of this remarkable plant. The path ahead will require not only scientific rigor but also open and informed social discussion about cannabis' place in our societies.

References